1. Department of Marine Microbiology and Biogeochemistry, NIOZ Royal Netherlands Institute for Sea Research, University of Utrecht, Texel, The Netherlands
2. Department of Freshwater and Marine Ecology, Institute for Biodiversity and Ecosystem Dynamics (IBED), University of Amsterdam, Amsterdam, The Netherlands
3. Ocean Biogeochemistry & Ecosystems Research Group, National Oceanography Centre, Southampton, UK
4. British Antarctic Survey, Natural Environmental Research Council, Cambridge, UK
5. Institute of Aquaculture, University of Stirling, Stirling, Scotland, UK