1. Southern Seas Ecology Laboratories, School of Biological Sciences and the Environment Institute, University of Adelaide, Adelaide, SA 5005, Australia
2. Primary Industries and Regions South Australia, Fisheries and Aquaculture, 25 Grenfell Street, Adelaide, SA 5000, Australia
3. Australian Research Council Centre of Excellence for Coral Reef Studies, School of Biological Sciences, The University of Queensland, St Lucia, QLD 4072, Australia