1. Fisheries & Aquaculture Science (Cefas), Centre for Environment, Pakefield Road, Lowestoft NR33 0HT, UK
2. Fisheries & Aquaculture Science (Cefas), Centre for Environment, 55 Briseham Road, Brixham TQ5 9NX, UK
3. University of Newcastle, 40-42 Great North Road, Newcastle upon Tyne NE1 7RU, UK