1. Institute of Oceanography, National Taiwan University , Taipei 10617 , Taiwan
2. NOAA National Marine Fisheries Service, Pacific Islands Fisheries Science Center , 1845 Wasp Boulevard, Building 176, Honolulu, HI 96818 , United States
3. Fisheries Resources Institutes, Japan Fisheries Research and Education Agency , 2-12-4, Fukuura, Kanazawa, Yokohma, Kanagawa 236-8648 , Japan
4. School of Aquatic and Fishery Science, University of Washington , 1122 NE Boat Street, Seattle, WA 98195 , United States