1. Bren School of Environmental Science and Management, Marine Science Institute, University of California, Santa Barbara, CA 93106, USA
2. National Oceanic and Atmospheric Administration, National Marine Fisheries Service, Alaska Fisheries Science Centre, 7600 Sand point Way NE, Seattle, WA 98115, USA
3. School of Aquatic and Fishery Sciences, University of Washington, 1122 Boat Street, Seattle, WA 98195, USA