1. Swedish University of Agricultural Sciences, Department of Aquatic Resources, Institute for Freshwater Research, Stångholmsvägen 2, SE-17893 Drottningholm, Sweden
2. ONEMA, pôle gest'aqua, 65 rue de Saint Brieuc, 35042 Rennes Cedex, France
3. INRA, U3E (1036), pôle gest'aqua, 65 rue de Saint Brieuc, 35042 Rennes Cedex, France