1. The University of Sydney, School of Life and Environmental Sciences, Camperdown, NSW 2006, Australia
2. The Sydney Institute of Marine Science, Mosman, NSW 2088, Australia
3. The University of New South Wales, School of Biological, Earth and Environmental Sciences, Kensington, NSW 2052, Australia
4. New South Wales Department of Primary Industries, Port Stephens Fisheries Institute, Taylors Beach, NSW 2316, Australia