1. School of Aquatic and Fishery Sciences, University of Washington, Seattle, WA 98250, USA
2. NOAA Alaska Fisheries Science Center, 17109 Pt Lena Loop Rd, Juneau, AK 99801, USA
3. NOAA Alaska Fisheries Science Center, 7600 Sand Point Way NE, Seattle, WA 98115, USA
4. NOAA Pacific Marine Environmental Laboratory, 7600 Sand Point Way NE, Seattle, WA 98115, USA
5. Department of Ecology and Evolutionary Biology, Yale University, New Haven, CT 06511, USA