1. Department of Ecology and Evolutionary Biology, Princeton University, NJ 08544, USA
2. National Oceanic and Atmospheric Administration, Northwest Fisheries Science Center, Seattle, WA 98112, USA
3. School of Marine Sciences, University of Maine, ME 04469, USA
4. Global Economic Dynamics and the Biosphere Program, The Royal Swedish Academy of Sciences, Stockholm 10405, Sweden
5. Oregon State University, College of Earth, Ocean and Atmospheric Sciences, OR 97331, USA