1. Departamento de Biologia Animal and cE3c, Faculdade de Ciências, Centre for Ecology, Evolution and Environmental Changes, Universidade de Lisboa, Bloco C2, Campo Grande, Lisboa 1749-016, Portugal
2. MARE—Marine and Environmental Sciences Centre, ISPA—Instituto Universitário, Rua Jardim do Tabaco 34, Lisboa 1149-041, Portugal
3. Department of Aquatic Resources, Swedish University of Agricultural Sciences, Havsfiskelaboratoriet, Turistgatan 5, Lysekil 453 30, Sweden