1. Institute for Marine and Atmospheric Research Utrecht (IMAU), Department of Physics, Utrecht University , Utrecht, Utrecht, 3584 CC , The Netherlands
2. Centre for Complex Systems Studies, Utrecht University , Utrecht, Utrecht, 3584 CE , The Netherlands
3. Oceanic Fisheries Programme, Pacific Community , Noumea 98848 , New Caledonia
4. Centre for Conservation Ecology and Genomics, Institute for Applied Ecology, University of Canberra , Australian Capital Territory , Australia