1. Coastal and Marine Resources Centre, ERI, University College Cork, Glucksman Marine Facility, Irish Naval Base, Haulbowline, Cobh, Cork, Ireland
2. School of Biological, Earth and Environmental Sciences, University College Cork, Distillery Fields, North Mall, Cork, Ireland
3. Fisheries Science Services, Marine Institute, Rinville, Oranmore, Galway, Ireland
4. Department of Pure and Applied Ecology, Institute of Environmental Sustainability, Swansea University, Swansea SA2 8PP, UK