1. School of Aquatic and Fisheries Sciences (SAFS) and Cooperative Institute for Climate, Ocean and Ecosystem Studies (CICOES), University of Washington , Seattle, WA 98036 , United States
2. Antarctic Ecosystem Research Division, NOAA Southwest Fisheries Science Center , La Jolla, CA 92037 , United States
3. Institute of Marine Research , Bergen N-5817 , Norway
4. Aqualyd Ltd., 341 Higgins Road, RD 1 , Wakefield 7095 , New Zealand