1. Lyell Centre, Institute for Life and Earth Sciences, Heriot-Watt University , Edinburgh EH14 EHS , UK
2. Environment and Sustainability Institute and Centre for Ecology and Conservation, University of Exeter , Penryn Campus, Penryn, Cornwall TR10 9FE , UK
3. Ocean Futures Research Cluster, School of Science, Technology & Engineering, University of the Sunshine Coast , Maroochydore, QLD 4556 , Australia
4. Department of Coastal Systems, NIOZ Royal Institute for Sea Research , PO Box 59, 1790 AB, Texel , The Netherlands
5. British Antarctic Survey, Natural Environment Research Council, High Cross , Madingley Road, Cambridge CB3 0ET , UK