1. Organismal and Evolutionary Biology Research Program, Faculty of Biological and Environmental Sciences, University of Helsinki , PO Box 56, FI-00014 Helsinki , Finland
2. Biodiversity Unit, University of Turku , FI-20014 Turku , Finland
3. Norwegian Institute for Nature Research (NINA) , NO-7485 Trondheim , Norway
4. Marine Scotland Science, Freshwater Fisheries Laboratory , Faskally, Pitlochry PH16 5LB , UK
5. Faroe Marine Research Institute , Nóatún 1, FO-110 Tórshavn , Faroe Islands
6. Natural Resources Institute Finland (Luke) , POB 413, FI-90014 Oulu , Finland
7. Department of Arctic and Marine Biology, UiT The Arctic University of Norway , NO-9037 Tromsø , Norway