1. Université de Caen Normandie , UMR7208 BOREA, 14000 Caen , France
2. France Énergies Marines , 29280 Plouzane , France
3. LEMAR Plouzane , 29280 Plouzane , France
4. GEOMAR Helmholtz Centre for Ocean Research Kiel, Marine Ecology Research Division Kiel , 24148 Kiel, DE , Germany
5. National Research Council of Italy, Institute of Biosciences and Bioresources , 50019 Sesto Fiorentino, Firenz, IT , Italy