1. Department of Integrative Biology, 4575 SW Research Way, Oregon State University , Corvallis, OR 97333 , USA
2. School of Biological Sciences, Washington State University , Pullman, WA 99164 , USA
3. Department of Biology, Whitman College , Walla Walla, WA 99362 , USA
4. Center for Reproductive Biology, Washington State University , Pullman, WA 99164 , USA