1. School of Plant Sciences, Haramaya University, Dire Dawa, Ethiopia
2. Department of Zoology and Entomology, University of Fort Hare, Alice, South Africa
3. Department of Crop Production, University of Swaziland, Luyengo Campus, Swaziland
4. Department of Biological and Environmental Sciences, Walter Sisulu University, Mthatha, South Africa
5. Department of Biological Sciences and Biotechnology, Botswana International University of Science and Technology, Private Bag 16, Palapye, Botswana
6. IRD/CNRS, UMR IRD 247 EGCE, Laboratoire Evolution Génomes Comportement et Ecologie, Avenue de la terrasse, BP1, 91198, Gif-sur-Yvette, France
7. Université Paris-Sud, Orsay, France