1. State Key Laboratory of Tree Genetics and Breeding, Key Laboratory of Tree Breeding and Cultivation of State Forestry Administration, Research Institute of Forestry, Chinese Academy of Forestry, No.1 Dongxiaofu, Xiangshan Road, Beijing 100091, China
2. Key Laboratory of Horticulture Science for Southern Mountains Regions of Ministry of Education, College of Horticulture and Landscape Architecture, Southwest University, No.2 Tiansheng Road, Chongqing 400715, China
3. State Key Laboratory of Tree Genetics and Breeding, Northeast Forestry University, No.26 Hexing Road, Harbin 150040, China
4. Department of Biology, Centre for Forest Biology, University of Victoria, 3800 Finnerty Road, Victoria BC, V8P 5C2, Canada