1. Department of Plant and Environmental Sciences, Edisto Research and Education Center, Clemson University , Blackville, SC , USA
2. USDA-APHIS-PPQ-Science & Technology, Identification Technology Program , Fort Collins, CO , USA
3. Department of Plant and Environmental Sciences, Pee Dee Research and Education Center, Clemson University , Florence, SC , USA
4. Department of Biological Sciences, Clemson University , Clemson, SC , USA