1. Division of Neurology/Molecular Brain Science, Kobe University Graduate School of Medicine, Kobe 650-0017, Japan
2. Division of Cell Physiology, Kobe University Graduate School of Medicine, Kobe 650-0017, Japan
3. Laboratory of Animal Resources, Center for Disease Biology and Integrative Medicine, The University of Tokyo, Tokyo 113-0033, Japan
4. Department of Neurology, Graduate School of Medicine, The University of Tokyo, Tokyo 113-0033, Japan