1. School of Food Engineering and Biotechnology, Tianjin University of Science and Technology, Tianjin 300457, P. R. China
2. Tianjin Institute of Industrial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, P. R. China
3. Key Laboratory of Systems Microbial Biotechnology, Chinese Academy of Sciences, Tianjin 300308, P. R. China