1. Graduate School of Agriculture, Kyoto University , Sakyo-ku, Kyoto 606–8502, Japan
2. Mushroom division, National Institute of Horticultural and Herbal Science , Rural Development Administration, Bisan-ro, Eumseong-gun, Chungcheongbuk-do, 22709, Republic of Korea
3. Institute of Edible Fungi, Shanghai Academy of Agricultural Sciences , Shanghai 201403, China