1. School of Public Health, Lanzhou University , 199 Donggang West Road, Lanzhou, Gansu Province 730000 , China
2. Central South Inventory and Planning Institute of National Forestry and Grassland Administration , 143 Xiangzhang East Road, Changsha, Hunan Province 410014 , China
3. State Key Laboratory of Grassland Agro-Ecosystems, Center for Grassland Microbiome, College of Pastoral Agriculture Science and Technology, Lanzhou University , 768 Jiayuguan West Road, Lanzhou, Gansu Province 730020 , China
4. Institute for Tropical Biodiversity and Sustainable Development, University Malaysia Terengganu , Kuala Nerus, Terengganu 21030 , Malaysia