1. School of Environmental Science, The University of Shiga Prefecture , 2500 Hassaka-cho, Hikone, Shiga 522-8533 , Japan
2. Graduate School of Agriculture, Kyoto University, Kitashirakawa oiwakecho , Sakyo-ku, Kyoto 606-8502 , Japan
3. Gifu Prefectural Research Institute for Forests , 1128-1 Sodai, Mino, Gifu 501-3714 , Japan
4. Kitami Institute of Technology , 165 Koen-cho, Kitami, Hokkaido 090-8507 , Japan