1. Department of Biological Sciences, Wayne State University , Detroit, Michigan , USA
2. Center of Molecular Medicine and Genetics, Wayne State University School of Medicine , Detroit, Michigan , USA
3. Department of Molecular and Human Genetics, Human Genome Sequencing Center, Baylor College of Medicine , Houston, Texas , USA
4. Evolutionary Biology of Invertebrates, Institute of Evolution and Ecology, University of Tübingen , Germany
5. Department of Ophthalmological, Visual, and Anatomical Sciences, Wayne State University School of Medicine , Detroit, Michigan , USA