1. Institute of Deep-sea Science and Engineering, Chinese Academy of Sciences , Sanya 572000 , China
2. State Key Laboratory of Freshwater Ecology and Biotechnology, Institute of Hydrobiology, Chinese Academy of Sciences , Wuhan 430072 , China
3. Academy of Plateau Science and Sustainability, Qinghai Normal University , Xining , China
4. University of Chinese Academy of Sciences , Beijing 100049 , China
5. Center for Excellence in Animal Evolution and Genetics, Chinese Academy of Sciences , Kunming, 650223 , China