1. Department of Ecology, Biodiversity and Animal Behaviour, Te Aka Mātuatua/School of Science, University of Waikato , Hamilton , New Zealand
2. Division of Ecology and Evolution, Research School of Biology, Australian National University , Canberra, ACT , Australia
3. Commonwealth Scientific and Industrial Research Organisation, Land & Water, Black Mountain Laboratories , Canberra, ACT , Australia
4. National Research Collections Australia, Commonwealth Scientific and Industrial Research Organisation , Black Mountain, Canberra, ACT , Australia