1. Department of Biology and Angelo State Natural History Collections, Angelo State University, ASU Station 10890, San Angelo, TX 76909-0890, USA
2. U.S. Geological Survey, Alaska Science Center, Anchorage, AK 99508, USA
3. Department of Environment, Government of Yukon, Whitehorse, YT, Y1A 2C6, Canada
4. Wildlife Research and Management, 3680 Non Road, Fairbanks, AK 99709, USA
5. Department of Biology and Museum of Southwestern Biology, University of New Mexico, Albuquerque, NM 87131, USA