1. Haub School of Environment and Natural Resources, University of Wyoming , Laramie, Wyoming 82072 , USA
2. Department of Zoology and Physiology, University of Wyoming , Laramie, Wyoming 82071 , USA
3. USDA Forest Service, Rocky Mountain Research Station , Missoula, Montana 59801 , USA
4. Wyoming Game and Fish Department , Lander, Wyoming 82520 , USA