1. Department of Wildlife, Fisheries and Aquaculture, Mississippi State University , Mississippi State, MS 39762 , United States
2. The School of Natural Resources, University of Tennessee , Knoxville, TN 37996 , United States
3. Mississippi Veterinary Research and Diagnostic Laboratory, College of Veterinary Medicine, Mississippi State University , Pearl, MS 39208 , United States
4. United States Department of Agriculture, Animal and Plant Health Inspection Service, Wildlife Services, National Wildlife Research Center , Fort Collins, CO 80521 , United States