1. Department of Wildlife Ecology and Conservation, University of Florida , Gainesville, Florida 32611 , USA
2. Sanibel-Captiva Conservation Foundation , 3333 Sanibel-Captiva Road, Sanibel, Florida 33957 , USA
3. J.N. Ding Darling National Wildlife Refuge, U.S. Fish and Wildlife Service , Sanibel, Florida 33957 , USA