1. Mammal Research Institute, Department of Zoology and Entomology, University of Pretoria, Pretoria, 0002, South Africa
2. Centre for Invasion Biology, Eugène Marais Chair of Wildlife Management, University of Pretoria, Pretoria, 0002, South Africa
3. Department of Botany and Zoology, Stellenbosch University, Stellenbosch, 7602, South Africa
4. Department of Life and Consumer Sciences, University of South Africa, Roodepoort, 1710, South Africa