1. Division of Wildlife Conservation, Alaska Department of Fish and Game , Anchorage, AK 99518 , United States
2. Department of Environmental and Life Sciences, Trent University , Peterborough, ON K9L 0G2 , Canada
3. Program on the Environment, Department of Natural Sciences, University of Alaska Southeast , Juneau, AK 99801 , United States
4. Department of Geography, University of Victoria , Victoria, BC V8W 2Y2 , Canada
5. Division of Wildlife Conservation, Alaska Department of Fish and Game , Juneau, AK 99811 , United States