1. Ecology Department, Montana State University, P.O. Box 173460, Bozeman, MT 59717-3460, USA
2. Montana Fish, Wildlife and Parks, 1420 E 6th Avenue, Helena, MT 59620, USA
3. Land Resources and Environmental Sciences Department, Montana State University, 334 Leon Johnson Hall, P.O. Box 173120, Bozeman, MT 59717-3120, USA