1. Faculty of Agriculture, Tokyo University of Agriculture and Technology , Fuchu, Tokyo 183-8509 , Japan
2. Institute of Global Innovation Research, Tokyo University of Agriculture and Technology , Fuchu, Tokyo , 183-8509 , Japan
3. United Graduate School of Agricultural Science, Tokyo University of Agriculture and Technology , Fuchu, Tokyo , 183-8509 , Japan
4. Faculty of Regional Environmental Science, Tokyo University of Agriculture , Setagaya, Tokyo 156-8502 , Japan
5. Illinois Natural History Survey, University of Illinois , Champaign, Illinois , 61820 , USA