1. Museum of Vertebrate Zoology , Department of Integrative Biology, University of California, Berkeley, Berkeley, California 94720 , USA
2. Biology Department, Mills College at Northeastern University , Oakland, California 94613 , USA
3. Museum of Vertebrate Zoology, Department of Integrative Biology, University of California , Berkeley, Berkeley, California 94720 , USA
4. Department of Biology, University of Wisconsin–Eau Claire , Eau Claire, Wisconsin 54702 , USA