1. Key Laboratory of Adaptation and Evolution of Plateau Biota, Northwest Institute of Plateau Biology, Chinese Academy of Sciences , Xining 810001 , China
2. University of Chinese Academy of Sciences , Beijing 100049 , China
3. School of Geography Science, Qinghai Normal University , Xining 810016 , China
4. Agricultural Service Center of Maduo County , Guoluo 813500 , China
5. Qinghai Key Laboratory of Animal Ecological Genomics, Northwest Institute of Plateau Biology, Chinese Academy of Sciences , Xining 810001 , China