1. Conservation Department, Minnesota Zoo, Apple Valley, MN, USA
2. Department of Fisheries, Wildlife & Conservation Biology, University of Minnesota, St. Paul, MN, USA
3. Natural Resources Research Institute and Department of Biology, University of Minnesota, Duluth, MN, USA
4. Wildlife Research Unit, Minnesota Department of Natural Resources, St. Paul, MN, USA
5. School of Environmental and Forest Sciences, University of Washington, Seattle, WA, USA
6. Wildlife Research Unit, Minnesota Department of Natural Resources, Forest Lake, MN, USA
7. Department of Veterinary Population Medicine, College of Veterinary Medicine, University of Minnesota, St. Paul, MN, USA