1. Nature Conservation and Landscape Ecology, University of Freiburg, 79106 Freiburg, Germany
2. Biodiversity Conservation and Management, University for Development Studies, PO.Box TL1882, Tamale, Ghana
3. Senckenberg Biodiversity and Climate Research Center, 60325 Frankfurt, Germany
4. Department of Theoretical and Applied Biology, College of Science, Kwame Nkrumah University of Science and Technology, PMB Kumasi, Ghana
5. Department of Biology, Conservation Ecology, University of Marburg, 35043 Marburg, Germany