1. Centre for Genomic & Experimental Medicine, Institute of Genetics & Cancer, University of Edinburgh, Western General Hospital , Edinburgh EH4 2XU , UK
2. Cancer Research UK Scotland Centre, Institute of Genetics & Cancer, University of Edinburgh, Western General Hospital , Edinburgh EH4 2XU , UK
3. Medical Research Council Human Genetics Unit, Institute of Genetics & Cancer, University of Edinburgh, Western General Hospital , Edinburgh EH4 2XU , UK
4. Department of Biomedicine, Aarhus University , 8000 Aarhus , Denmark