1. Wolfson Centre for Age-related Diseases, King’s College London, Guy’s Campus , London SE1 1UL , UK
2. Molecular Neuropharmacology and Genetics Laboratory, Department of Neuroscience, Faculty of Health and Medical Sciences, University of Copenhagen , 2200 Copenhagen , Denmark
3. Pain Centre Versus Arthritis, School of Life Sciences, University of Nottingham , Nottingham NG5 1PB , UK