1. Department of Medicine, Sulpizio Cardiovascular Center, University of California San Diego , 9452 Medical Center Drive, La Jolla, San Diego, CA 92037-7411 , USA
2. BHF Centre for Cardiovascular Science, University of Edinburgh , Edinburgh , UK
3. Usher Institute, University of Edinburgh , Edinburgh , UK
4. Cardiovascular Research Institute Basel (CRIB) and Department of Cardiology, University Hospital Basel, University of Basel , Basel , Switzerland