1. Systems Immunity Research Institute, School of Medicine, Cardiff University , Cardiff CF14 4XN , UK
2. Babraham Institute, Babraham Research Campus , Cambridge CB22 3AT , UK
3. Department of Pharmacology, Department of Chemistry and Biochemistry, University of California , San Diego , La Jolla , CA 92093-0601 , USA
4. Department of Bioengineering, University of California , San Diego , 9500 Gilman Drive , La Jolla , CA 92037 , USA
5. Swansea University Medical School , Singleton Park, Swansea SA2 8PP , Wales , UK
6. Boehringer Ingelheim Espana SA, Carrer de Prat de la Riba , 50, 08174 Sant Cugat del Vallès, Barcelona , Spain