1. Signalling Research Centres BIOSS and CIBSS, University of Freiburg , Schänzlestr. 18, 79104 , Freiburg , Germany
2. Institute of Biology II, Faculty of Biology, University of Freiburg , Schänzlestr. 1, 79104 , Freiburg , Germany
3. Spemann Graduate School of Biology and Medicine (SGBM), University of Freiburg , Albertstr. 19A, 79104 , Freiburg , Germany
4. Cell Biology and Biophysics Unit, European Molecular Biology Laboratory , Meyerhofstraße 1, 69117 , Heidelberg , Germany
5. Institute of Biology III, Faculty of Biology, University of Freiburg , Schänzlestr. 1, 79104 , Freiburg , Germany