1. Department of Applied Biological Chemistry, Graduate School of Agricultural and Life Sciences, University of Tokyo, Tokyo 113–8657, Japan
2. Department of Preventive Intervention for Psychiatric Disorders, National Institute of Mental Health, National Center of Neurology and Psychiatry, Tokyo 187–8553, Japan
3. ERATO Touhara Chemosensory Signal Project, JST, University of Tokyo, Tokyo 113–8657, Japan
4. WPI International Research Center for Neurointelligence, University of Tokyo Institutes for Advanced Study, Tokyo 113-0033, Japan