1. School of Forestry, Northern Arizona University , 200 E Pine Knoll Drive, Flagstaff, AZ 86011 , USA
2. USDA Forest Service, Northern Research Station , 202 Anheuser-Busch Natural Resources Building, Columbia, MO 65211 , USA
3. USDA Forest Service, Forest Health Protection , Arizona Zone, 2500 S Pine Knoll Drive, Flagstaff, AZ 86001 , USA
4. Department of Biological Sciences, Northern Arizona University , 610 S Knoles Drive, Flagstaff, AZ 86011 , USA