1. Graduate School of Life and Environmental Science, Shimane University , 1060 Nishikawatsu-Cho, Matsue, Shimane, Japan
2. Graduate School of Natural Science and Technology, Shimane University , 1060 Nishikawatsu-Cho, Matsue, Shimane, Japan
3. Department of Life Sciences, Faculty of Life and Environmental Sciences, Shimane University , 1060 Nishikawatsu-Cho, Matsue, Shimane, Japan
4. Estuary Research Center, Shimane University , 1060 Nishikawatsu-Cho, Matsue, Shimane, Japan
5. Interdisciplinary Center for Science Research, Shimane University , 1060 Nishikawatsu-Cho, Matsue, Shimane, Japan
6. Institute of Agricultural and Life Sciences, Academic Assembly, Shimane University , 1060 Nishikawatsu cho, Matsue, Shimane, Japan