1. Graduate School of Science, Tohoku University , Aoba-ku, Sendai , Japan
2. Graduate School of Life Science, Tohoku University , Aoba-ku, Sendai , Japan
3. Graduate School of Science and Engineering, Teikyo University , Utsunomiya, Tochigi Japan
4. Agro-Biotechnology Research Center, Graduate School of Agricultural and Life Sciences, The University of Tokyo , Bunkyo-ku, Tokyo , Japan