1. Faculty of Life and Environmental Sciences, University of Tsukuba, 1-1-1 Tennodai, Tsukuba, Ibarakim, Japan
2. Institute of Agriculture, Academic Assembly, Shinshu University, 8304, Minamiminowa, Nagano, Japan
3. Plant Productivity Systems Research Group, RIKEN Center for Sustainable Resource Science, 1-7-22, Suehiro, Tsurumi-ku, Yokohama, Japan
4. Department of Biological Sciences, Graduate School of Science, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo, Japan